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A researcher at Chicago’s Natural History Museum returns from South America with some crates containing his findings. When the crates arrive at the museum without the owner there appears to be very little inside. However, police discover gruesome murders on the cargo ship that brought the crates to the US and then another murder in the museum itself. Investigating the murders is Lt. Vincent D’Agosta who enlists the help of Dr. Margo Green at the museum - she has taken an interest in the contents of her colleague’s crates. Unknown to both there is a large creature roaming the museum which is gearing itself up for a benefit reception which the city’s mayor is to attend.

The sequel of the first movie, the New York Zoo Animals, Alex the Lion, Marty the Zebra, Melman the Giraffe and Gloria the Hippo, still stranded on Madagascar, started to leave the island. All of a sudden, they landed in the wilderness of Africa. There, Alex soon meet the rest of his family and starting to have trouble communicating with them after much of his time at the Central Park Zoo.

The year is 1936. A professor who studies archeology named Inidana Jones is venturing in the jungles in South America searching for a golden statue. Unfortunately, he sets off a deadly trap doing so, miraculously, he escapes. Then, Jones hears from a museum curator named Marcus Brody about a biblical artifact called The Ark of the Covenant, which can hold the key to humanly existence. Jones has to venture to vast places such as Nepal and Egypt to find this artifact. However, he will have to fight his enemy Renee Belloq and a band of Nazis in order to reach it.

At New York’s Central Park Zoo, a lion (Stiller), a zebra (Rock), a giraffe (Schwimmer), and a hippo (Smith) are best friends and stars of the show. But when one of the animals goes missing from their cage, the other three break free to look for him, only to find themselves reunited … on a ship en route to Africa. When their vessel is hijacked, however, the friends, who have all been raised in captivity, learn first-hand what life can be like in the wild.