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Taking a romantic tropical island week off with her boyfriend, an ambitious, decisive New York girl agrees to help her magazine out by covering a story on a neighbouring island. The only plane available is piloted by a laid-back heavy drinker with whom she shares a mutual dislike. But she has no choice and he can’t refuse the money. When the plane crash-lands on an uninhabited island with little chance of rescue they both wish they had made other arrangements - at least to start with.

At New York’s Central Park Zoo, a lion (Stiller), a zebra (Rock), a giraffe (Schwimmer), and a hippo (Smith) are best friends and stars of the show. But when one of the animals goes missing from their cage, the other three break free to look for him, only to find themselves reunited … on a ship en route to Africa. When their vessel is hijacked, however, the friends, who have all been raised in captivity, learn first-hand what life can be like in the wild.