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Security guard Larry Daley infiltrates the Smithsonian Institute in order to rescue Jedediah and Octavius, who have been shipped to the museum by mistake.

When the coach of the France soccer team is killed by a poisoned dart in the stadium in the end of a game, and his expensive and huge ring with the diamond Pink Panther disappears, the ambitious Chief Insp. Dreyfus (Kevin Kline) assigns the worst police inspector Jacques Clouseau (Steve Martin) to the case. His intention is to give a diversion to the press, while he uses his best men to chase the killer and thief. He assigns Gendarme Gilbert Ponton (Jean Reno) to work with Closeau and inform each step of the investigation. When Clouseau is nominated with honor to the highest prize in France, Dreyfus decides to humiliate Clouseau and take him out of the case. However Clouseau has already solved the mystery.

In New York, the unemployed divorced Larry Daley is a complete loser. His son Nick is very disappointed with his father that is gong to be evicted, and Larry accepts the job of night watchman in the Museum of Natural History as the substituted for three old security guards that have just retired to raise some money and pay his bills. In his first nightshift, Larry realizes that everything at the museum comes to life at night. The Museum transforms in a complete chaos with the inexperienced Larry, and he learns that since an old Egyptian stone came to the Museum in 1950, the was statues comes to life until dawn. When Larry brings his son to spend a night with him, the three old guards break in the Museum to stole the magic stone. Larry organizes the historic characters to help him to arrest the criminals and save the museum.